actinopterygii adaptation american english animal betta burdigala betta chini betta chloropharynx betta hipposideros betta livida betta macrostoma betta miniopinna betta pallifina betta persephone betta simplex betta smaragdina betta spilotogena betta tomi biological classification bubble nest chordate common name evolution fiber fishbase freshwater genus gourami greek alphabet iucn red list inch indonesia integrated taxonomic information system malaysia mouthbrooder open directory project pellet penang betta phosphorus pieter bleeker protein siamese fighting fish singapore species spotted betta thailand type species united nations environment programme united states valenciennes wikimedia commons world conservation monitoring centre