african bush elephant african forest elephant african savanna elephant african elephant anancus asian elephant borneo elephant columbian mammoth cuvieronius dwarf elephant elephas elephas recki gomphothere gomphotherium hypselephas indian elephant john godfrey saxe mammuthus imperator mammuthus meridionalis mammuthus subplanifrons mammuthus sungari moeritherium north african elephant palaeoloxodon palaeomastodon paratetralophodon population genetics primelephas pygmy mammoth sri lankan elephant stegodon stegomastodon stegotetrabelodon steppe mammoth sumatran elephant syrian elephant tetralophodon woolly mammoth zygolophodon